English: A complex bout emitted by a male of P. septemradiatus . Left panel: video recording; the fish, which is a resident territorial owner, is oriented towards the cage containing the intruder (on the right, not visible). The oscillogram (top right panel) and spectrogram (bottom right panel) of the emitted bout show a train of pulses followed by a tonal segment. Acoustical and video recordings are synchronised. Note the rapid downward movements of the head made during the pulse emissions, and the behaviours preceding and following the vocalisation (dorsal fins' display and jump, respectively). During the sequence, the mudskipper is also retracting its eyes into the dermal cups positioned below the orbits (“blinking”), to clean and moisten the eye surface while out of water.